nigrospora fungus black center syndrome banana

Late last week, the Colombian Agricultural Institute (ICA) in Bogot confirmed that four plantations in northern Colombia have been quarantined because of suspected infection with Fusarium wilt tropical race 4 (TR4), a fungus that kills plants by clogging their vascular system. Nigrospora is a fungal disease that causes the centre of the banana to turn dark red. If a banana is mistreated in the shipping process by being allowed to get cold or has too much weight put on top of it. banana black center syndrome. Even though nigrospora doesnt look particularly appealing, it is not actually harmful to humans. 3). Scientists have sequenced the genomes of three fungal diseases that currently threaten banana crops, and found something disturbing - the fungi have evolved to the point where they could wipe out the most popular banana crops in five to 10 years. Black Center Syndrome in bananas is caused by a fungus called Nigrospora fungus. i.e. Conidia are single-celled, smooth, spherical to ovoid, and 1420 m in diameter. However, red spots or red streaks may be a sign of another closely-related bacterial disease. Many recipes for banana cakes, bread, or cookies call for overripe bananas. Eating rotten bananas can affect the human body like many other foodborne illnesses. I decided to start this blog to share my recipes and knowledge with you and would love for you to join me. Heat to a boil, then boil 2 to 3 minutes. Russ Caid, special director of banana and technical services for Chiquita, said that the condition is called black center syndrome. Despite our best efforts, sometimes it is impossible to avoid the blackened centers of our bananas. He explained that once bananas have begun to ripen, they are very fragile and must be handled with care. However, once these bananas are closer to their final destination, either at distribution centers or the supermarket, handling may become less careful, causing black center syndrome. Banana producers insist that if your banana has a blackened core due to being dropped or hit at the wrong time, the fruit is still safe to eat. (Explained for All Kind of Nuts! Bananas that have blackened in the middle, for the most part, are not safe to eat. He explained that once bananas have begun to ripen, they are very fragile and must be handled with care. He explained that once bananas have begun to ripen, they are very fragile and must be handled with care. Black center syndrome can be an indication of two things: As bananas ripen, their skin starts to change color from yellow to brown and finally to black. Jordan's line about intimate parties in The Great Gatsby? The fungus won't make you physically sick if you accidentally eat it. For this reason, you should always check the banana before eating them. Nigrospora is a fungal disease that causes the centre of the banana to turn dark red. International Banana Association, which reported that this was most likely due to adverse conditions at some point after picking. They are inedible. For decades, biotech researchers and conventional breeders were foiled in their efforts to bring disease resistance to the Cavendish [banana] or to hybridize a replacement for the thick-skinned,. Is it safe to eat a banana with black center Syndrome? Nigrospora is a fungal disease that causes the centre of the banana to turn dark red. Applications of super-mathematics to non-super mathematics. (All You Need to Know), How to Salt Unsalted Nuts? But, the fungal infection can look reddish-brown instead of black. Such bananas always come with a mushy feel and can get highly soft. According to an article on Foodandfizz.com, " Black Center Syndrome in bananas is caused by a fungus called Nigrospora fungus. Since Nigrospora is mainly a plant pathogen that rarely causes opportunistic infections in humans, data on treating drugs is scarce. Black Center Syndrome There are really two reasons that a banana can look ripe and good to eat, but it will have a black core on the inside. Why did the Soviets not shoot down US spy satellites during the Cold War? Mokillo, moko, and blood disease bacteria are bacterial diseases that can cause red discoloration in bananas as well. Septate hyaline hyphae, hyaline or slightly pigmented conidiophores, and conidia are visualized. Bananas are a delicious fruit with many health benefits that it can provide; it is used in entrees, desserts and can be peeled and eaten by itself. Black center syndrome can be an indication of two things: A fungal infection Mishandling at the distribution center As bananas ripen, their skin starts to change color from yellow to brown and finally to black. What does the blackened core of a banana taste like? What Is Black Center Syndrome in Bananas? But, if it is of concern to you, simply throw the banana away. We received a more thorough explanation from Chiquita Bananas along with the admission that the problem is occurring with more frequency than ever before. 1). BBS usually occurs when bananas are stored improperly. So, as you can see, nigrospora is hardly something for humans to worry about. Jim Melban of the California Dry Bean Association said that although he can`t say point blank why your beans aren`t cooking evenly, he can suggest some reasons. Banana growers and shippers are aware of that and insulate the bananas from any rough handling. I've eaten bananas in Thailand and. An eight-year-old girl bit into a banana to be greeted with a red streak inside. its nigrospora fungus, black center syndrome is more from bruising and other ailments of the banana could be fungal ailments as well but the isolated hardness of the center means its a banana infected with nigrospora. How to make banana bread moist after baking. Nigrospora species produce glass-like, septate hyphae that become darkly pigmented with age. Offices in Canada are the most affected by the Basidiospores mold group?! During the ripening process, bananas skin turns yellow, brown, and black. That being said, there have only ever been 4 reported cases of nigrospora-related eye or skin infections. That is one sign you could look for, but the fungus can still look black. The blackened center is usually very hard and crunchy, making it an unpleasant surprise when you bite into your breakfast banana, expecting it to be soft and sweet. The patient was managed medically and fully recovered after the therapy [8]. Thankfully, this type of fungus doesnt result in lasting harm, with most discomfort being psychosomatic, so Dionne made a full recovery and was discharged from the hospital. Scientists Have Linked Selfies To Narcissism, Addiction And Mental Illness, Essential Oils And Brain Injuries: Heres What You Are Not Being Told, Rare Monkeypox Virus Has Started Reappearing, Heres What You Need To Know, Heart Surgeon Speaks Out About The True Cause Of Heart Disease, Kind FedEx Driver Shovels Snow Off Womans Porch When He Learns Her Husband Died, Man Builds A Tiny Blue House At Narrow Alley As Revenge To Stop Neighbors From Using It, I Dont Want To Live Anymore And I Want To Die. Read This First. A black center is also an indication of mishandling, such as being dropped or crushed under heavy crates. This is common knowledge amongst those in the banana industry, and precautions are taken to ensure that the bananas delivered do not arrive with ugly black centers. Can the Spiritual Weapon spell be used as cover? Bananas with this issue may appear yellow from the outside, but they are, in fact, black on the inside. Guess Who Her Dad Is. Ear rot diseases have been observed and stalk rot diseases are becoming increasingly common. We throw them away. The cold will turn the banana skin brown, but your fruit will be okay to eat. Infrequently, this bacteria enters into the banana through wind, water, or birds and produces an effect . If you go for example to Asia and try one of the local sorts, you may well run into fruits with lots of seeds. If the blackened part is only the center, you may think the outside is still safe to eat. EMN-150715-132938001. Spread occurs through vascular bundles, which become discoloured brown or dark red, and finally purplish or black. Rotten bananas smell awful, and you will be able to smell the difference between ripe and rot. Which average weight % of the banana is edible? Nigrospora is differentiated from Humicola by its very black conidia that originate from hyaline, inflated conidiophores. Have you noticed this? Often, this black center is a type of fungus called Nigrospora. Yellow and brown bananas are safe to eat. Frequently Asked Questions About Banana Black Center Syndrome. Each person will have to decide what to do with their banana if they find a blackened core. Bananasare one of the most popular fruits today. Not until you peel the banana and take a bite will you see the black center. Nigrospora is prevalent in tropical climates, where bananas are grown, and will also infect other tropical fruits. Looked forward to eating it. Beyond the visual clues you can get when checking your bananas, you will also be able to smell whether or not a banana is rotten. However, if your banana smells bad, then it probably tastes just as bad. Either way, it doesnt look particularly appetizing, but is it something you should be concerned about? There is a possibility that you may suffer from diarrhea and vomiting if you eat contaminated food that has a fungal infection. Site design / logo 2023 Stack Exchange Inc; user contributions licensed under CC BY-SA. If the banana is still yellowish-white, then the fruit is still good. Nigrospora is a fungal infection that turns the banana's center dark red. She was later diagnosed with a fungal corneal ulcer, which showed the presence of nigrospora. Are carrots safe to eat after they have turned black? If the bananas are moldy, smell rotten, or have a black center aka Read More Is It Ok To Eat Bananas With Black In The Middle? Nigrospora can infect the fruit in tropical climates where . Is it safe to eat? DONT EAT. There are a variety of plant diseases that can cause the inside of bananas to take on a red discolouration. This banana had a bit of bruising I wasn't too concerned until I noticed the core (and only the core) was dark brown/black in color. And, while neither will kill you, I dont relish eating a fungal banana. So today, I peeled one. So, you may ask: Is it safe to eat if the banana is black in the middle? Diet and Nutrition. The diseases are: 1. In one such case, a healthy female in India developed an eye infection and a corneal ulcer after an injury caused by a cows tail containing soil and plant matter contaminated with spores of Nigrospora. While unattractive to eat, these diseases do not pose a health risk to humans; however, if in doubt, toss it out or compost it. This will cause the center of a banana to turn dark red. That is Nirgispora, a banana disease that hasn't been documented to harm humans, but we . (Explained for All Kind of Nuts! It does not mean the banana was overripe or rotten. Ethylene is a ripening hormone that is naturally present in ripe fruits. But if the piece of fruit involved is in a bad condition, it can affect someones health for the worst. (The Fascinating Answer! On the outside, bananas with this issue may appear yellow, but they are black inside. Conclusion About Banana Black Center Syndrome, How to Prevent Grease Buildup in the Kitchen. The insect getting in them. The trick is to eat them or cook with them before they go completely black. We received a more thorough explanation from Chiquita Bananas along with the admission that the problem is occurring with more frequency than ever before. In fact, some said that almost all the bananas they had purchased lately had such an unpleasant, dark center that they were inedible. According to Melban, the hot soak method not only increases the beans` ability to absorb water during cooking, it also removes about 75 percent of the undigestible sugars from the beans that are responsible for causing gas. Scientific Publishers, India. Has Microsoft lowered its Windows 11 eligibility criteria? Also, if they are not ripening fast enough for you, perhaps try to buy yellower bananas rather than green ones. https://www.youtube.com/watch?v=ex0URF-hWj4, The Terrifying Truth About Bananas (https://www.youtube.com/watch?v=ex0URF-hWj4). She puts the meat in a microwave-safe colander placed over a glass bowl, then cooks it on high power in 2- to 3-minute increments until it is done. Re: Help - I found this in my banana. Bananas grow in warm places, but they can also withstand low temperatures of up to 14 degrees. Ouch! Why are non-Western countries siding with China in the UN. The conidiogenous cells on the conidiophores are inflated, swollen, and ampulliform in shape. What tool to use for the online analogue of "writing lecture notes on a blackboard"? rev2023.3.1.43269. Even a small drop can cause the centers to go black. Good luck, and hopefully, all your bananas in the future do not have blackened centers. Mumbai's busy Sion Hospital has reported 24 cases of the fungal infection in the past two months, up from six cases a year, according to Dr Renuka . Studies have shown that soaking beans in several different sodium solutions increases their ability to absorb water during cooking. FoodandFizz 2022 |Iseli Media Group | Privacy Policy | About us | Contact | Terms&Conditions | Mediavine Trellis Child on Trellis Framework by Mediavine, Frequently Asked Questions About Core of My Banana is Black. FoodandFizz 2022 |Iseli Media Group | Privacy Policy | About us | Contact | Terms&Conditions | Mediavine Trellis Child on Trellis Framework by Mediavine, 4 Things You Need to Know About Banana Black Center Syndrome, 1. Here is the method recommended by the California Dry Bean Association for soaking 1 pound of dry beans: Put 1 pound of beans in a large pot with 10 cups of water and 2 teaspoons salt, if desired. Sometimes, Black Center Syndrome produces a hard, crunchy banana core that is colored black or dark brown. So, there is room for confusion about when a banana is ok to eat and when it isnt. However, while blackened center bananas pose no deadly threat, the black parts indicate that something went wrong somewhere. In the case of bananas, if they start turning dark brown, it is not advisable to consume them since, as we have discussed, decay will soon start to set in. News broke late last week that a fungal disease that kills Cavendish banana plants has been detected in Latin America for the first time. Of the bunch, a couple started to ripen much earlier than the rest. The fact that the fruit looks completely healthy and delicious is one of the more disturbing elements of Black Center Syndrome. Meredith, D. S. (1961). And for all the health nuts in the house (I know I am), bananas are an excellent source of: If you want to grow a banana tree in your backyard (yes, it is possible), first you have to live in a place that is not very cold. People who have fungal toenail infections often have a fungal skin infection on the foot, especially between the toes (commonly called athlete's foot, ringworm on the foot, or tinea pedis). Even so, there are a couple of well-documented cases of nigrospora in humans. I know this is old but for anybody reading here for info in the future Black Center Syndrome is Not the Same as Overripe Bananas. So the consumer is grossed out, the vendors are unhappy, and the producers are also frustrated. Therefore, it can cause black centers in some bananas, rendering them unfit for consumption. However, there are some things to consider before you do so to avoid further complications. So, if your banana has a black core, it is better not to eat it because it is likely rotten. Among the most common and economically important Fusarium species associated with . shivaay film full movie hd. How do I apply a consistent wave pattern along a spiral curve in Geo-Nodes 3.3? Get Special Gift: Industry-Standard Mold Removal Guidelines. If you notice red spots, red bruising, or red discoloration of a banana, this could also point to mokillo, moko, or blood disease bacterium. While watching for signs of rot and mold is essential, do not let a little character on your banana peel hold you back from enjoying it. Nigrospora sphaerica is the best-known species of the genus Nigrospora. Nigrospora can infect the fruit in tropical climates where bananas are grown. Mokillo, moko, and blood disease bacterium are bacterial diseases that can also cause red discoloration in bananas. Banana black center syndrome (BBS) is a condition where the banana turns black after being cut open. Posted on Last updated: November 25, 2022, How to Age Bananas for Banana Bread The 4 Best Methods. The genus Nigrospora belongs to the phylum Ascomycota and is comprised of typically dark-spored asexual, filamentous fungi (dematiaceous fungi). When light pressure is applied, the pulp transforms into a mushy liquid that squirts out from the base [6]. If your banana has a black center on an otherwise healthy piece of fruit, it may have a fungal infestation. Thus, there is no reason to eat something like that. Eat a banana, and you. December. The Fusarium fungus invades young roots or root bases, often through wounds. One of the important indicators is the fruit on the inside of the peel. I was recently making some banana bread, and as I was mashing the bananas up, one of them was all black on the inside. Fruits are healthy for people of all ages and are vital to a balanced diet. As a consequence, the older beans won`t cook quite the same way as newer ones. The banana tree is not a tree but an herb. And, even if the outside of the fruit looks ripe, if the core is black, dont eat it. Several weeks ago, this column responded to a reader`s question about bananas, which, on the outside, looked bright yellow and perfect but upon peeling revealed dark slimy centers. ), Whats the Difference Between Banana Chips and Dehydrated Bananas? Merlin rushed to check on the bananas that they had, and when cut open, the bananas revealed dark red streaks down their centers. To be safe, make sure that your banana is a safe one to eat. You can always scrape out the yellow parts whenever you have a banana with a black core since they are still safe to eat. It's important to scout for stalk rot diseases now to determine which fields are at greatest risk of lodging and should be harvested first. However, it is not thought to be deadly for humans. Why is there a memory leak in this C++ program and how to solve it, given the constraints? This fungus is actually red, but in bananas, it looks brown or is so dark it looks black. However, when you select bananas, know when you are going to eat them. So, if a box falls off the pallet or something heavy is thrown on top of the bananas in shipping, the bananas could have Black Center Syndrome. Of course, its arguable for many people, especially since throwing a banana seems wasteful. Depending on isolates, sporulation may take more than three weeks [4]. Nigrospora sphaerica is an airborne filamentous fungus in the phylum Ascomycota.It is found in soil, air, and plants as a leaf pathogen. The most common reason your banana has red in it is due to the fungal disease nigrospora. As The Whole Portion confirms, these are bananas to avoid. If you buy green bananas, be careful not to drop them or cause them any physical injuries and stress. And finally, I recommend not eating bananas with this syndrome, but I leave that to each of you to make an informed decision. Banana growers and shippers are aware of that and insulate the bananas from any rough handling. The black core may also be caused by mishandling bananas in shipping. With that being said, mokillo often referred to as red finger, is rarely found in bananas. The nail may also become separated from the nail bed. I just do not want to take the risk of my banana carrying the fungus. But, in reality, as we now know, any illness was potentially more in her mind than anything actually physical. However, if the fruit is browning or has brown spots, it is probably still good. N. sphaerica, as well as N. oryzae, N. sachari, and N. musae, are known for causing squirter disease on banana plants. Mc Ginnis, M. R. (1980). It is best not to eat this black core because it could be a fungal infection. An unusual deep fungal infection with. Bananas with this syndrome generally look fresh and ripe with white flesh on the fruit, but there is a black core that can sometimes be crunchy when bitten into. As the bananas ripen, they get sweeter. Laboratory analysis confirmed that lesions were primarily caused by N. spherica and secondary infections were caused by a resistant Staphylococcus aureus strain. Chicagoans have another story to tell. It happens because of the oxidation of the banana's natural sugars. Also, have we said how scary a black center banana looks? However, its pathogenic role as a causative agent is not well-known [1847, 2218]. How Many Scoville Is Buffalo Wild Wings Mango Habanero Sauce. Or brown centers like yours. Asexual spores (conidia) develop at the tips of translucent conidiophores that rise at a 90 angle to the vegetative hyphae. A fungus can set in. Conclusion Only the Core of my Banana is Black, How to Prevent Grease Buildup in the Kitchen. Yes, it is safe to eat a banana with black in the middle. What causes this and is there a way to prevent it from happening? Is it safe to eat a black banana in the middle? December 17, 2021; translational research vs qualitative research; Sporulation of N. sphaerica causes its initial white coloured colonies to rapidly turn black. Introduction. Once it's infected a plantation, its fruit is toast. That is each individuals right. Another aspect of bananas that sometimes turns people off is how easily the fruit bruises. Nigrospora is a filamentous dematiaceous fungus widely distributed in soil, decaying plants, and seeds. A large banana has 121 calories. All fruit is good to eat as long as it doesn't smell or taste funny so just use your senses. Fungal nail infections may cause nails to become discolored, thick, fragile, or cracked. A strain of the Fusarium fungus, which causes so-called . When it comes to bananas, when they start to turn dark brown, it may not be best to consume them, because as we have mentioned above, decay is beginning to set in. How Many Scoville Is Buffalo Wild Wings Mango Habanero Sauce. Merlin rushed to check on the bananas that they had, and when cut open, the bananas revealed dark red streaks down their centers. I had to learn to cook out of necessity - but its become a hobby that I (and my family) enjoy immensely. As the bananas age, the flavor becomes even more potent. Also known as Fusarium fungus, the blight blackens bananas from the inside out. Allen, R. N. (1970). If so, then you may want to discard the whole fruit completely. The symptoms initially begin by the formation of a dark core or a broken line of dark-red rubbery material along the center core of the banana fruit or near its end (Fig. Download the industry-standard guidelines that Mold Busters use in their own mold removal services, including news, tips and special offers: document.getElementById( "ak_js_1" ).setAttribute( "value", ( new Date() ).getTime() ); Written by:Jelena SomborskiMycologistMold Busters, Edited by:Dusan SadikovicMycologist MSc, PhDMold Busters. Sheath rot Nectria foliicola Mycosphaerella musicola Pseudocercospora musae Sooty mold Limacinula tenuis: Speckle Mycosphaerella musae: Squirter (black end disease) Nigrospora sphaerica: Stem-end rot Colletotrichum musae: Trachysphaera finger rot Connect and share knowledge within a single location that is structured and easy to search. Are Bananas with Black Center Syndrome Safe to Eat? But just because banana is bruised does not mean it is not edible. Steps to Ensure Your Bananas Do Not Get Black Center Syndrome. It is said that the experience left her feeling sick, so after a non-emergency phone consultation, her mother took her to Accident and Emergency. Yi-Ming Fan, et al (42) expressed Nigrospora as a rarely opportunistic pathogen in humans. Just dont eat the black parts. Bananas are slightly radioactive! Commenters on Merlins Facebook post reasoned that it was not possible to check every single piece of fruit in the grocery store. While unappealing to eat, these diseases affecting bananas are not a threat to human health, however when in doubt, throw it out or compost them. -. In addition, it's somewhat dry and hollow, unlike normal banana flesh. Then, I would brush my teeth and gargle mouthwash for a while. Realistically, nigrospora cannot be considered to be a true human pathogen. Nigrospora can infect the fruit in tropical climates where bananas are grown. The sexual stage of Nigrospora species belongs to the genus Khuskia [1]. The black center syndrome affects only the center of the banana and does not spread to the rest of it. This is not true; if your banana is showing signs of age, that does not mean that the fruit is bad on the inside. 1990 ), and submitted into GenBank with accession number MW228165. Nigrospora has been isolated from cutaneous lesions of a leukemic patient and from a case with keratitis. Nigrospora can cause a dark center in the banana which can eventually lead to a softening of the entire banana pulp. Caid said that at Chiquita they are very concerned about the problem and are trying to orient and train people at all levels of handling to eliminate the condition. Symptoms included swelling, redness, and vision problems (Fig. If the bananas are moldy, smell rotten, or have a black center aka black center syndrome, a fungal condition do not eat them. None of these diseases are actually harmful to humans. Nigrospora Disease of Rice: The disease of rice has widespread occurrence in the rice-growing tracts of the world. For this reason, its safe to assume that you can eat the outer parts without worries as long as you discard the black core. Nigrospora is a fungal disease that causes the centre of the banana to turn dark red. However, you should only eat the outer parts and not the ones with black color in them. Brown Spot of Rice. Still, many people find the red appearance unappealing, which makes them queasy. Hi, my name is Daniel and I am passionate about cooking. One can simply cut away the injured portion of the fruit and then continue to enjoy the deliciousness. Although these experiments can`t be duplicated at home because they used several different types of sodium, it has been found that sodium chloride (table salt) can be effective. Remove from the heat and soak 4 hours. Cubense (Foc). Can a black banana in the middle make you sick? Some may even opt to try to salvage the ripe parts. However, you should discard the black center since its inedible and may cause several problems if consumed. That looks to be. In contrast, the disease has not been proven to be toxic to humans. The blackened core is a sign of Black Center Syndrome. Skip to content Franais Mold Library Our Services Locations Resources About Us Contact Us Search for:Search Search Absidia AcremoniumMenu Toggle It is a syndrome, a pejorative, and an acronym of our times: NIMBY, or Not In My Back Yard. Instead of browning ground beef on the stove and draining away as much excess fat as possible, she cooks it in the microwave oven. Most fruits, when opened, will turn black. This fungus is actually red, but in bananas, it looks brown or is so dark it looks black. It is a common laboratory contaminant. Often, this black center is a type of fungus called Nigrospora. If you find that your banana has red in it, this is likely to be the fungal disease nigrospora. When Is A Banana Too Ripe? Nigrospora is characterized as presenting micronematous or semimacronematous conidiophores, which are branched, flexuous, hyaline to brown, smooth, and usually reduced to conidiogenous cells;. As the bananas ripen, there comes a particular time when the fruit will blacken in the center if mishandled. One common misconception that many have with bananas is that they must be perfectly yellow; otherwise, they are not worth eating. The first, as mentioned above, is theNigrosporafungus sickens the fruit. Bananas, harvested from my garden. How do I know if a black banana is too old to be eaten? She then took to Google to see what the streaks meant, and quickly discovered that they were riddled with the fungus Nigrospora. Unfortunately, he concluded, there is no way to tell if a banana has black center syndrome until it is peeled. Vinegar or lime added to bananas can slow down the ripening process. Part of the challenge is that they can look extremely similar in bananas. The black center syndrome causes this black center, and the good thing is that it only affects the center and does not spread towards the outer parts. Bananas with this syndrome generally look fresh and ripe with white flesh on the fruit, but there is a black core that can sometimes be crunchy when bitten into. My date has some black powder in it. This fungal disease, known as Squirter's (because eventually the soft. Mango, banana, papaya, pineapple, and avocado are categorized as major tropical fruits grown for local consumption, export, and sources of income to the growers. Yellow and brown bananas are safe to eat. According to Kenny, 1 pound of beef labelled 80 percent lean yielded slightly over 3 ounces of fat when it was cooked this way. Only several cases of human infections caused by Nigrospora species have been reported so far, referring to respiratory allergic reactions as well as skin, nail, and eye infections [8, 9]. They concluded that the incidence was extremely rare. One explanation was that perhaps they had been held at too low a temperature. See the summary of synonyms and teleomorph-anamorph relations for Nigrospora spp. Yes, a banana with a black center or middle can make you sick, but only when eating the blackened parts. N. oryzae is also a causative agent of ear rot and comb rot of maize, while N. sphaerica is famous for causing the squirter disease on bananas [3, 7]. Upon further investigation and DNA testing, it was found that the onychomycosis was caused by nigrospora.

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